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No vehicle in World of Tanks can operate without a complete virtual crew. The crew operates the vehicle under the direction of the player. The size of a complete crew varies, depending on the vehicle in question - In-game crews presently range from a minimum of 2 to a maximum of 6 crew members. Crew members fill one of five designated roles, also known as their Major Qualification within a vehicle. The 5 roles are Commander, Gunner, Driver, Radio Operator, and Loader. All tanks must have these roles covered; in some cases, an individual crew member covers more than one role, in which case they would have Multiple Major Qualifications.

Just as vehicles accumulate experience for every battle, so do crew members. Similar to the Service Record for a player or vehicle, each crew member has a Personal File, which records his own battle statistics, medals, and primary (Major Qualification) Training Level, as well as the levels of any Skills and/or Perks he has acquired. Crew members can receive a headstart on their primary Training Level by receiving training purchased with credits or gold. Note that this kind of training only affects a crew member's Training Level in his Major Qualification, and as such, cannot be used to give a headstart or boost to Skills/Perks training.

The degree of Proficiency that each crew member achieves through training directly affects and improves the performance of their vehicle in battle. If a crew member is wounded or killed in battle (both have identical effects), the vehicle's performance will suffer, specifically in the area affected by his role. Secondary skills/perks that give a slight bonus to certain aspects of the vehicle's performance will also be disabled, and thus there can also be slight loss in performance to areas of the tank that are outside of the knocked-out crew member's specific Major Qualification.

A knocked out Driver, for example, would reduce the acceleration, speed, and maneuverability of your vehicle.

If all crew members are knocked out, the vehicle will become inoperable.